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When we compared gene relationships among terrestrial and aquatic ecosystems, we found that terrestrial studies exhibited significant positive correlations between the abundance of genes and processes (r P n whereas those from aquatic studies did not (r P n This finding may reflect inherent differences between these habitats, due to a more rapid temporal decoupling of organisms and biogeochemical reactants in aquatic habitats where the matrix is more temporally static than soils. Within aquatic and terrestrial biomes, we also examined more specific habitat types and found that agricultural systems and coastal aquatic systems exhibited a significant positive gene relationship ((r P n and (r P n respectively), whereas other habitat types (forest, grassland, meadow, tundra, estuary and lake; Figure 2) did not show a significant relationship. The strength of the correlation between agricultural and coastal studies may be due to strong signal to noise ratios associated with fertilizer at agricultural sites, which due to well documented runoff and connectivity may also impact coastal ecosystems.
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